托福TPO2第一篇文章的原文：The Origins of Cetaceans(鲸类起源)。
TPO-2-1：The Origins of Cetaceans
It should be obvious that cetaceans—whales, porpoises, and dolphins—are mammals. They breathe through lungs, not through gills, and give birth to live young. Their streamlined bodies, the absence of hind legs, and the presence of a fluke1 and blowhole2 cannot disguise their affinities with land dwelling mammals. However, unlike the cases of sea otters and pinnipeds (seals, sea lions, and walruses, whose limbs are functional both on land and at sea), it is not easy to envision what the first whales looked like. Extinct but already fully marine cetaceans are known from the fossil record. How was the gap between a walking mammal and a swimming whale bridged? Missing until recently were fossils clearly intermediate, or transitional, between land mammals and cetaceans.
Very exciting discoveries have finally allowed scientists to reconstruct the most likely origins of cetaceans. In 1979, a team looking for fossils in northern Pakistan found what proved to be the oldest fossil whale. The fossil was officially named Pakicetus in honor of the country where the discovery was made. Pakicetus was found embedded in rocks formed from river deposits that were 52 million years old. The river that formed these deposits was actually not far from an ancient ocean known as the Tethys Sea.
The fossil consists of a complete skull of an archaeocyte, an extinct group of ancestors of modern cetaceans. Although limited to a skull, the Pakicetus fossil provides precious details on the origins of cetaceans. The skull is cetacean-like but its jawbones lack the enlarged space that is filled with fat or oil and used for receiving underwater sound in modern whales. Pakicetus probably detected sound through the ear opening as in land mammals. The skull also lacks a blowhole, another cetacean adaptation for diving. Other features, however, show experts that Pakicetus is a transitional form between a group of extinct flesh-eating mammals, the mesonychids, and cetaceans. It has been suggested that Pakicetus fed on fish in shallow water and was not yet adapted for life in the open ocean. It probably bred and gave birth on land.
Another major discovery was made in Egypt in 1989. Several skeletons of another early whale, Basilosaurus, were found in sediments left by the Tethys Sea and now exposed in the Sahara desert. This whale lived around 40 million years ago, 12 million years after Pakicetus. Many incomplete skeletons were found but they included, for the first time in an archaeocyte, a complete hind leg that features a foot with three tiny toes. Such legs would have been far too small to have supported the 50-foot-long Basilosaurus on land. Basilosaurus was undoubtedly a fully marine whale with possibly nonfunctional, or vestigial, hind legs.
An even more exciting find was reported in 1994, also from Pakistan. The now extinct whale Ambulocetus natans ("the walking whale that swam") lived in the Tethys Sea 49 million years ago. It lived around 3 million years after Pakicetus but 9 million before Basilosaurus. The fossil luckily includes a good portion of the hind legs. The legs were strong and ended in long feet very much like those of a modern pinniped. The legs were certainly functional both on land and at sea. The whale retained a tail and lacked a fluke, the major means of locomotion in modern cetaceans. The structure of the backbone shows, however, that Ambulocetus swam like modern whales by moving the rear portion of its body up and down, even though a fluke was missing. The large hind legs were used for propulsion in water. On land, where it probably bred and gave birth, Ambulocetus may have moved around very much like a modern sea lion. It was undoubtedly a whale that linked life on land with life at sea.
1994年，巴基斯坦报道了一个更令人兴奋的发现。目前已经灭绝的鲸鱼Ambulocetus natans(可以步行的鲸类)4900万年前曾在古地中海生活过。比Pakicetus晚大约300万年，比Basilosaurus早900万年左右。幸运的是，被发现的Ambulocetus natans保留着完整的后肢。它的后肢很强壮，底部有长足，非常像现在的鳍足类动物。这些后肢使得他们既能在陆地行走又能在海里游行。虽然Ambulocetus natans保留了尾巴，但它们缺少现代水生鲸类动物用于行动的主要身体部位——尾片。不过，从Ambulocetus的脊椎结构上可以看出，即使缺少尾片，它们也能像现代鲸鱼那样通过身体背部的上下摆动来游走。大的后肢通常被当作是水中前行的发动机。在它们可能交配繁殖的陆地上，Ambulocetus行动起来非常像现代海狮。毫无疑问，鲸鱼是连接着陆地生命和海洋生命的物种。
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